Herpetology of Ethiopia & Eritrea


Ptychadenidae Dubois, 1987



Hildebrandtia macrotympanum (Boulenger, 1912)


Description:
A medium sized frog, up to 55 mm. Vomerine teeth in two oblique series between the choanae, close to the inner borders of the latter. Head feebly depressed, much broader than long; snout rounded, as long as the orbit ;canthus rostralis obtuse, loreal region concave ; nostril a little nearer the eye than the end of the snout ; interorbital space nearly as broad as the upper eyelid; tympanum distinct, larger than the eye. Fingers extremely short, blunt, first and second equal; toes short, blunt, half webbed; subarticular tubercles small; a large, compressed, very prominent but not sharp-edged inner metatarsal tubercle, its length equalling that of the inner toe. Tarso-metatarsal articulation reaching the tympanum; tibia two-fiths the length of head and body. Skin smooth. Pale greyish brown above, with dark brown dots, and a dark brown band extending from the end of the snout, through the nostril and eye, to the groin, where it breaks up into spots, and expanding into a large blotch below the eye and on the temple; a brown line borders the upper lip; lower lip with large brown blotches; lower parts white, with a few brown dots on the throat [according to BOULENGER].

Distribution:
It occurs from northern Somalia to southern Kenya and south-east Ethiopia.

Note:
Species of dry savannas and semi-arid areas, and it is probable, that activates only in the short rainy season. Breeds in periodic ponds.

Description original:
Boulenger, G. A. 1912. Descriptions of new African batrachians preserved in the British Museum. Annals and Magazine of Natural History, Series 8, 10: 140-142.

Described as Pyxicephalus macrotympanum

Common english name: Somali Ornate Frog

Hildebrandtia macrotympanum - Filtu, Harerge keflehager, Ethiopia



Hildbrandtia ornata (Peters, 1878)


Diagnosis:
A medium sized frog, with typical Y-shaped pattern on throat, and with characteristic dorsal pattern. The body plump, blunt snout. Males have paired lateral vocal sacs whose slits are situated immediately anterior to the armpit. The large tympanum is set apart from the eye and is barely visible on young animals, but conspicuous on adults. The tympanum reaches the size of 0.5 to 0.6 diameter of the eye. The hind legs are rather short. Maximal length up to 65 mm, Sometimes shown even 70 mm SVL.
Typical color on the left picture is from Gambella region.

Distribution:
Species with a very large area of distribution in tropical and subtropical parts of Africa, from Senegal and Gambia east to Kenya and Ethiopia and south to northern Namibia, northern Botswana, southern and northern Mozambique and South Africa. Not recorded in Guinea-Bissau, Guinea, Burkina Faso, Niger, Chad and Sudan, but probably occur in these countries.
The finds from Ethiopia originate from my collections, it is not always reported for Ethiopia (Largen,Spawls).

Behavior:
The species inhabits the Sudan savanna, it apparently prefers rather dry and open landscapes. Presumably distributed over all savanna regions south of the Sahara. Eggs measured in diameter 1.4 mm. Individual eggs are deposited in the shallow water, they are often scattered on a large surface. Reproductive behavior starts immediately after the rainy season, in the time when the savannah flooded with water. Tadpoles are large and muscular, with long tails. Base of the tail is particularly wide. A marked green metallic basic coloration is usually discernible, and this metallic glimmer can be considered as a feature which distinguishes these tadpoles from those of any other species.. The metamorphosis does not last longer than four weeks. Tadpoles are carnivorous and feed on aquatic insects and other tadpoles of the same species. Metamorphosed frogs are 20mm long.

Description original:
Peters, W. C. H. 1878. Über die von Hrn. J. M. Hidebrandt während seiner letzten ostafrikansichen Reise gesammelten Säugethiere und Amphibien. Monatsberichte der Königlichen Preussische Akademie des Wissenschaften zu Berlin 1878: 194-209.

Described as Pyxicephalus ornatus

Common english name: African Ornate Frog

Hildbrandtia ornata - Gambella, Illubabor keflehager, Ethiopia


Lanzarana largeni (Lanza, 1978)

Diagnosis:
Omosternum widely forked posteriorly; outer metatarsal tubercle present; inner metatarsal tubercle well developed but not compressed; terminal phalanges rounded; main webbing always reaching middle subarticular tubercle of 4th toe, at least during the breeding season; throat immaculate.

Distribution:
This species is endemic to Somalia. Most records are from the south of the country in Bakool and Bay Provinces. It has also been recorded near the coast, north of Mogadishu, at Bud Bud in Galguduud Province, and near Garoowe in Nugaal Province. Recorded localities are the general vicinity of Garoowe, Bud Bud, El Ure (Uegit), Sciù Manàs (Baidoa), Dinsor, Rahole, Afgoi, and Showli Berdi (Bardera). It has been recorded up to 500m. It is possible, that it occurs in adjacent areas of Ethiopia, in Juba and Wabe Shebelle basins.

Description original:
Lanza, 1978, Monit. Zool. Ital., N.S., Suppl., 10: 234

Common english name: Lanza's Frog



Ptychadena anchietae (Bocage, 1868)

Ptychadena anchietae, Baro Gambella hotel, Gambella town, W Ethiopia-©Trailin

Diagnosis:
A medium-sized frog reach up to 50 mm, with a pointed nose and a characteristic color pattern with black mask-like stripes present on the face, dark spots on the flanks and a brown and yellow marbled pattern on the backs of the thighs. The dorsum is grayish olive brown, and the belly is white or yellowish white. Some individuals have darker blotches scattered over the dorsum.

Distribution:
This species occur from Eritrea, Ethiopia, Djibouti and Somalia, south through East Africa to South Africa and Botswana, and west to Angola and southern Republic of the Congo. In Ethiopia the most abundant species of genus "Ptychadena" in lowland area.

Behavior:
It is a species of open country in woodland, savannah, grassland, and agricultural and suburban areas, but also found in forest clearings. It breeds in shallow temporary ponds. Small eggs, gray on top and white below, are laid in a single layer that floats on the surface of the water. Tadpoles are brown above and light below with a clear fin and may reach 45 mm in length.

Description original:
Bocage, J. V. B. d. 1868 "1867". Batraciens nouveaux de lAfrique occidentale (Loanda et Benguella). Proceedings of the Zoological Society of London 1867: 843-846.

Common english name: Anchieta's Ridged Frog

Ptychadena anchietae - Gewane, Wello keflehager, Ethiopia



Ptychadena cooperi (Parker, 1930)


Diagnosis:
A medium sized frog. SVL 45 mm. The head is broader than long. The upper lip is white or greyish. The snout is prominent and rounded terminally. The back has longitudinal rows of short, very indistinct or well defined, glandular folds and is brownish grey with small indistinct dark spots, and a fine white mid-dorsal stripe from snout to vent. The underside is smooth, and coloration is white with pale brown reticulation. A fold is present across the chest. The hind limbs have indistinct crossbars. The hinder side of thighs is dark, with numerous small, round, white spots. Small vocal sacs show externally, through short slits close to the angle of the mouth, but not extending backwards beyond it.
Vomerine teeth absent. Head broader than long ; snout prominent, rounded terminally, once and one-third the diameter of the eye ; canthus rostralis obtuse; loreal region very oblique, not concave ; nostril midway between eye and end of snout ; interorbital space as broad as the upper eyelid; tympanum distinct, 3/5 the diameter of the eye. First finger slightly shorter than the second, which is shorter than the fourth ; third slightly longer than the snout. Toes half-webbed, the third slightly longer than the fifth; a small inner, but no outer, metatarsal tubercle; a slight tarsal fold, no tarsal tubercle. Tibio-tarsal articulation reaching the anterior border of the eye. Tibia three times as long as broad, its length contained once and four-fifths in the distance from snout to vent. Back with longitudinal rows of short, very indistinct, glandular folds ; a short glandular ridge from the angle of the mouth to the front of the shoulder. Smooth beneath; a fold across the chest.
Colour in alcohol: Brownish grey above, with small indistinct darker spots; a dark streak from the end of the snout through the nostril and eye, broadening behind the latter to inelnde the whole of the tympanunr. A fine white mid-dorsal stripe from snout to vent; upper lip white. Hind limbs with indistinct crossbars ; hinder side of thighs dark, with numerous small, round, white spots. White beneath, closely reticulated with pale brown. Vocal sacs present, small, showing externally through short slits close to the angle of the mouth, but not extending backwards beyond it. Length from snout to vent 45 mm.; fore limb 25 mm ; hind limb 80 mm [according to PARKER].

Distribution:
This species is endemic to the highlands of central Ethiopia west of the Rift Valley, and southern Ethiopia east of the Rift Valley

Note:
This species is distributed in montane grassland and occasionally forest-edge habitats at 2,500-3,100 m. It is often found in urban habitats, such as near Addis Ababa. Breeds in permanent pools, temporary rain-fed pools and flooded fields.

Description original:
Parker, H. W. 1930. Report on the Amphibia collected by Mr. J. Omer-Cooper in Ethiopia. Proceedings of the Zoological Society of London 1930: 1-6.

Common english name: Dark-bellied Grass Frog




Ptychadena erlangeri (Ahl, 1924)


Diagnosis:
A medium sized frog, size of males 35 - 47 mm, females are larger 40 - 54 mm (Largen). Dorsum brown, brown-gray, with dark spots on the back. A similar to species P.neumanii.

Distribution:
This species occurs in the Rift Valley in Ethiopia, and in the highlands on both sides of the Rift Valley. It is generally found at 1,500-2,500 m asl, though it possibly extends as low as 1,300 m asl near the shore of Lake Abaya.

Note:
Inhabits localities with long grasses and similar coarse herbaceous vegetation in forest clearings, not necessarily very close to permanent water. Its breeding behaviour is unknown, but it presumably takes place in pools in forest clearings and at forest margins.

Description original:
Ahl, E. 1924. Über eine Froschsammlung aus Nordost-Afrika und Arabien. Mitteilungen aus dem Zoologischen Museum in Berlin 11: 1-12.

Common english name: Erlanger's Grass Frog

Ptychadena erlangeri - locality, Bonga, SW Ethiopia



Ptychadena filwoha Largen, 1997


Diagnosis:
A rather small frog. Males reach 35 mm, females 40 mm. The dorsum is green, grey-green, yellowish, or brown, with pale yellowish vertebral stripe. The belly is white or cream, the throat usually freckled.

Distribution:
P.filwoha is known only from two nearby locaties between 800-1000 m.n/m in north Rift Valley in Ethiopia (Awash - Hot springs, and river Awash). It is possible that he lives elsewhere in similar habitat in the region.

Behavior:
This species is associated with permanent water (in which it breeds), which is provided at one site by pools deep amongst lava boulders, and by effluent streams from a hot spring at the other site, in a sub-desert steppe of volcanic rubble, xerophilous grasses and dense Acacia thickets (Largen 2001). Neighborhood is covered with xerophytic grasses, acacia thickets and palm trees (Hyphaene thebaica).

Note:
Provisional synonymy with Ptychadena nilotica by Zimkus, Lawson, Barej, Barratt, Channing, Dash, Dehling, Du Preez, Gehring, Greenbaum, Gvoždík, Harvey, Kielgast, Kusamba, Nagy, Pabijan, Penner, Rödel, Vences, and Lötters, 2016, Mol. Phylogenet. Evol., 106: 265. (However, there are two morphotypes of Grass Frogs in the same locality and habitat)
Filwoha means in Amharic generally "hot springs".

Description original:
Largen, M. J. 1997. Two new species of Ptychadena Boulenger, 1917 (Amphibia, Anura, Ranidae) from Ethiopia, with observations on other members of the genus recorded from this country and a tentative key for their identification. Tropical Zoology. Firenze 10: 223-246.

Common english name: Hot Springs Grass Frog

Ptychadena filwoha - locality, Awash NP, Fillwoha, Ethiopia



Ptychadena harenna Largen, 1997


Distribution:
This species is known only from the type locality at the foothills of the Bale Mountains in altitude 1550 m. It has been found in Podocarpus forest amongst herbaceous vegetation on the forest floor, and in a shallow, stagnant pool, devoid of aquatic vegetation, close to the edge of a fast-flowing river (Largen 2001).

Note:
It has been found in Podocarpus forest amongst herbaceous vegetation on the forest floor, and in a shallow, stagnant pool, devoid of aquatic vegetation, close to the edge of a fast-flowing river (Largen 2001). Unknown species.

Description original:
Largen, M. J. 1997. Two new species of Ptychadena Boulenger, 1917 (Amphibia, Anura, Ranidae) from Ethiopia, with observations on other members of the genus recorded from this country and a tentative key for their identification. Tropical Zoology. Firenze 10: 223-246.

Common english name: Harenna Forest Grass Frog




Ptychadena nilotica (Duméril and Bibron, 1841)


Popis:
Big "ranid" with pointy snout. SVL of males up to 57 mm, females 70 mm. Large, clearly visible tympanum. Two pairs of elongated dorsal ridges. On the lateral sides may present, in places, wart bumps. Males have paired vocal sacs. The tips of the fingers are not extended. Inner metatarsal callus is small and reaches only 0.2 to 0.4 of the length of the shortest finger. The belly skin is granulated. The colour is very variable. Dorsum mostly brown, or green-brown, often with wide brighter vertebral stripe, from tip of nose to the end of the body. The color of the stripe may be white, creamy, yellow, orange or green. Some individuals may miss this band. A few black spots may be present on the dorsal ridges. Dorsolateral ridges are bright. Brown tympanum has a bright center. Between the nostrils and eye is color dark. The vocal sacs are gray. Dark bands on the thighs are found only on the upper side. The upper lip is bright white. Belly white or yellowish. The inner parts of the legs are white and orange.

Distribution:
Nile drainage of Egypt and Sudan into Ethiopia, thence south into eastern Dem. Rep. Congo, Uganda, Rwanda, Burundi, western and southern Kenya and Tanzania, south to northern Botswana, Mozambique, Swaziland, and extreme eastern Rep. South Africa; presumably to be found in the Caprivi Strip of Namibia, Zimbabwe, Zambia, as well as northern Somalia, Djibouti, and southern Eritrea.

Behavior:
Very adaptable, euryvalent species. Females lay eggs in the number about 1,000 pieces with a diameter of 1.1 mm. Dorsum of the tadpoles with brown spots. The tail is relatively short. These frogs breed almost all year round, before and after the rainy season, except during the dry season. They breeds in ponds, roadside ditches, swamps and waterholes, but also in the larger lakes and slow flowing rivers. The frogs are active during day even night. They prey grasshoppers, beetles and aquatic invertebrates, but also small vertebrates (another frogs, small rodents...).

Note:
Leapfrogging into new territory: How Mascarene ridged frogs diversified across Africa and Madagascar to maintain their ecological niche

Description original:
Duméril, A. M. C., and G. Bibron . 1841. Erpétologie Genérale ou Histoire Naturelle Complète des Reptiles. Volume 8. Paris: Librarie Enclyclopedique de Roret.

Common english name: Nile Grass Frog

Ptychadena nilotica - habitat, Asayta, Danakil, NE Ethiopia




Ptychadena nana Perret, 1980      !

Distribution:
The endemic species Of Ethiopia. This species is currently known only from two areas: Dida'a, in the Arsi Mountains east of the Rift Valley, and from the Bale Mountains (Böhme and Rödder 2011) It is likely to occur more widely on the eastern side of the Ethiopian plateau (A. Mengistu and S. Loader pers. comms. June 2012).

Behavior:
It occurs in montane grassland and has been found near streams, in cattle grazing fields and in mosaics of cropland, trees, shrubs, grassland and other natural vegetation and in roadside pools in rural towns and villages (Mengistu 2012; A. Mengistu pers. comm. November 2012). Its occurrence in altered habitats suggests that it has a degree of tolerance to habitat disturbance, although probably not to more intense urbanization or to intensification of use of agricultural chemicals in fields (A. Mengistu pers. comm. November 2012). As with other congeners, it is thought to breed by larval development in water.

Description original:
Perret, J.-L. 1980. Sur quelques Ptychadena (Amphibia, Ranidae) d'Ethiopie. Monitore Zoologico Italiano. Nuova Serie, Supplemento. Firenze 13: 151-168.

Common english name: Smallest Grass Frog




Ptychadena neumanni (Ahl, 1924)


Diagnosis:
A medium sized frog. Males measure 30-40 mm, females 35-47 mm. Longitudinal ridges on the dorsum are more or less irregularly and extensively fragmented. The dorsum is light brown, yellowish or dark brown. On the back vertebral band, that is green, yellowisch, cream. The belly is cream, or white.

Distribution:
An endemic species of Ethiopian highlands. It occurs commonly on both sides of the Rift Valley, except the northernmost parts of the country. Range of altitude is 820-3,800 m.

Behavior:
It inhabits extremely wide range of habitats, including lowland grasslands, but also mountain forests, open grassland at moderate to high elevations, and even penetrates well into the Afro-alpine moorland zone. But intolerant of even mild aridity, and is conspicuously absent from all types of savannah. It breeds in large numbers in inundated land, as well as pools, streamlets and water-filled ditches, both temporary and permanent.

Description original:
Ahl, E. 1924. Über eine Froschsammlung aus Nordost-Afrika und Arabien. Mitteilungen aus dem Zoologischen Museum in Berlin 11: 1-12.

Common english name: Neumann's Grass Frog

Ptychadena neumanni - locality, Chebera Churchura, Kaffa keflehager, Ethiopia



Ptychadena porosissima (Steindachner, 1867)

Diagnosis:
A medium-sized ranid frog. Males reach 45 mm, and females 50 mm. The dorsum is brown, cream or grey. On the back darker spots or blotches, often with vertebral orange stripe.The belly is white or cream.

Distribution:
This species occurs from the east and the north to South Africa, in the west north to Angola, to the south of Democratic Republic of Congo, Rwanda, Uganda and Kenya. Isolated populations occur south of Lake Tana, in altitude up to 1900 m in Ethiopia. It was not recorded in Burundi, but is likely to occur there. Some individual records are presents in Kenya. It occurs especially in colder areas and is generally known from altitudes above 1200 m. Ethiopian locality is in the mountain pastures along the lake shores.

Behavior:
Adults feed opportunistically on terrestrial arthropods (Inger and Marx 1961), and Channing (2001) found prey to include earthworms, snails and other frogs. Males usually call from concealed positions amongst emergent vegetation within the water, supporting themselves on emergent plants, with the hind third of their bodies submerged (Passmore 1978). The breeding habitat includes vleis, inundated grassland and sedge pans.

Description original
Steindachner, F. 1867. Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Bafehlen des Commodore B. von Wüllerstorf-Urbair. Zologischer Theil. 1. Amphibien. Wien: K. K. Hof- und Staatsdruckerei.

Common english name: Striped Grass Frog




Ptychadena pumilio (Boulenger, 1920)

Diagnosis:
A small ranid with a moderately pointed snout. Adult males measure 25-32 mm, females 26-36 mm. Four pairs of symmetrically arranged longitudinal ridges on its back. The third pair, calculated from the spine, reaches only half the length of the body. All these ridges are continuous. On the flanks, some warts form a short ridge. The tympanum is clearly visible, reaching 0.7–0.9 of the eye diameter and appearing slightly concave. Males with paired lateral vocal sacs, enlarged thenar tubercles and swollen first fingers. Sturdy, moderately long hind limbs. The dorsum dark brown, olive, brown and green to bright green. The vertebral dark orange stripe may be present, but not always.

Distribution:
It occurs from Senegal to the northeast of Democratic Republic of Congo and to west Ethiopia in the east. In Ethiopia lives up to 1900 m.

Behavior:
A little known about way of life. The spawning sites are usually shallow ponds and ditches which are densely vegetated. During the dry season, this species is also encountered on the river banks. At least the borders of rainforest are inhabited, too. The habitats mentioned in the literature comprise savannas, grassland, humid and dry forests, gallery forests and the banks of all sorts of waters.

Description original:
Boulenger, G. A. 1920. Descriptions of three new frogs in the collection of the British Museum. Annals and Magazine of Natural History, Series 9, 6: 106-108.

Common english name: Dwarf Grass Frog




Ptychadena schillukorum (Werner, 1908)


Popis:
A medium-sized rather compact ranid with comparatively short and massive hind legs. The males reach up to 48 mm, females 50 mm. Males with paired lateral vocal sacs whose slits run parallel to the jaw border and end near the bases of the forearms, i.e. above the latter. The head width equals its length. The distance snout tip–nostril is less than the distance between the nostrils and the distance nostril–eye. The tympanum reaches 0.7 of the eye diameter. The dorsal and lateral ridges are divided into elongate wart-like structures. The skin of the flanks is granulated to warty. The foot is shorter than the shank, and the latter does not reach half of the SVL. The fourth toe with 1–2 free phalanges. The inner metatarsal tubercle is small, an outer one is absent (Stewart 1967, Poynton 1970). Very rarely light lines or bands on the back or on the dorsal part of the shanks. The rather discontinuous dorsal ridges are black. The light colored remains of the dorsolateral ridges have fine black borders. A black patch marks the eyelids. A dark "V" is present between the eyes. The anterior border of the vocal sac and the canthal area are somewhat darker, and the upper lip is light colored, showing no markings. This white color stretches to the upper arm where another isolated white bulge is found. The arms are feebly mottled. Black bars are present on the legs: three bars usually appear on the thighs, and 3–4 on the shanks. These bars are often paler centrally.

Distribution:
It occurs from Senegal east, to southwest Ethiopia, south Sudan, and to Somalia. South to Malawi, central Mozambique, to the southeast of the Democratic Republic of Congo and to Angola in the west.

Behavior:
A typical residents of savanna and semi-arid areas. Is linked to permanent water sources.
Males occupy during the rainy season flooded meadows, valleys and waterholes, where partially submerged in the water calls. Most often they are hiding in the flooded grass on the shore. Outside the rainy season will reside in dense vegetation at the edge of shallow ponds, swamps, irrigation channels and oxbow rivers. They hunt relatively large insects, cockroaches, butterflies, dragonflies and other arthropods.

Description original:
Werner, F. 1908 "1907". Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. Franz Werners nach dem agyptischen Sudan und Nord-Uganda. XII. Die Reptilien und Amphibien. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 116: 1823-1926.

Common english name: Schilluk Grass Frog

Ptychadena schillukorum - locality, Gamo Gofa keflehager, Ethiopia



Ptychadena tellinii (Peracca, 1904)

Ptychadena tellinii - Metemma, Ethiopia
Ptychadena tellinii - Metemma, Ethiopia

Diagnosis:
A medium-sized, somewhat stunted ranid with long hind legs. Adult males reach up to 40 mm, females 47 mm. Several rows of vaguely defined dorsal ridges, two of which stretch from the eyes to the center of the back where they gradually fade, two further ridges emerge anterior to the center of the back, stretching backward towards the vent, which they hardly ever touch. Besides, two pairs of very short ridges are usually located further outward. The tympanum is clearly visible. Kromě toho se dva páry velmi krátkých hřbenů nachází na vnější dorsolaterální straně. Tympanum je jasně viditelné a dosahuje 0,5 až 0,8 (obvykle 0,7) průměru oka. Males have paired lateral vocal sacs, enlarged thenar tubercles and swollen first fingers. Dorsum almost uniform red brown to red. Very rarely, a vaguely defined dark marking may be present in the center of the back. The red back is sharply separated from the belly by a dark band. Some animals have paler upper lips. The anterior parts of the male’s vocal sacs are black, the rest is pale beige. The webs are pigmented dark.

Distribution:
Benin, Burkina Faso, Cameroon, Central African Republic, Chad, Congo, the Democratic Republic of the, Eritrea, Ethiopia, Ghana, Mali, Nigeria, Senegal, Sierra Leone, Togo

Behavior:
In the dry season, is found both under stones near rivers and in dry savanna pools where it seeks refuge in crevices and under rotten wood. Gallery forests are never used as breeding sites. They are just crossed when the frogs move to the savanna, with the onset of the rains. As soon as the savanna ponds are filled with water, considerable choruses start to establish on the pond’s edges. Calling males either hide under vegetation, or they call from exposed sites on bare soil. The spawn is usually deposited in shallow water near the banks, but it floats occasionally to other sections of the pond.
We've found them in permanent residual pools along streams, where they lived together with freshwater crabs and fish.

Description original:
Peracca, M. G. 1904. Rettili ed Anfibi dell'Eritrea raccolti dal Dott. Achille Tellini nel 1903. Bollettino dei Musei di Zoologia e Anatomia Comparata della R. Universita di Torino 19 (467): 1-6.

Ptychadena tellinii - beside Metemma - Aykel road, Begemder keflehager, Ethiopia



Ptychadena wadei Largen, 2000

Permanent pond beside the road from Bahir Dar to Tiss Issat, approx 10 km away from the waterfalls.

Distribution:
This species is known only from a small area south-east of Lake Tana in Ethiopia, along the road from Bahar Dar to the Tisist Falls of the Blue Nile River, at 1,800-1,850m asl. It probably occurs more widely.

Note:
It lives in montane grassland and shrubland, and it has been found breeding in temporary pools and streamlets in montane grassland bordering the upper reaches of the Blue Nile River.

Description original:
Another new species of Ptychadena Boulenger 1917 from Ethiopia (Amphibia Anura Ranidae), MALCOLM J. LARGEN, Liverpool Museum, William Brown Street, Liverpool L3 8EN, England, Received 19 November 1999, accepted 23 March 2000























































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